The single-lens reflex (SLR) camera uses an automatic moving mirror system which permits the photographer to see exactly what will be captured by the film or digital imaging system, as opposed to non-SLR cameras where the view through the viewfinder could be significantly different from what was captured on film.

Prior to the development of SLR, all cameras with viewfinders had two optical light paths: one path through the lens to the film, and another path positioned slightly off to the side seen by the photographer. Because the viewfinder and the film lens cannot share the same optical path, the viewfinder is aimed to intersect with the film lens at a fixed point somewhere in front of the camera. This is more or less helpful for pictures taken at a middle or longer distance but a close-up shot framed in the viewfinder will not look the same as the film image. Moreover, focusing the lens of a non-SLR camera when it is opened to wider apertures (such as in low light or while using low-speed film) is not easy.

SLR cameras permit direct viewing using a pentaprism situated above the optical path through the lens to the film plane. Light is reflected by a movable mirror upwards into the pentaprism where it is reflected several times until it aligns with the viewfinder. When the shutter is released, the mirror moves out of the light path and the light shines directly onto the film, or in the case of a DSLR, the CCD or CMOS imaging sensor.

The focus can be adjusted manually by the photographer or automatically by the autofocus system. The viewfinder can include a matte focusing screen located just above the mirror system to diffuse the light. This system permits accurate viewing, composing and focusing, especially useful with interchangeable lenses.

Up until the 1990s, SLR was the most advanced photographic preview system available. But the recent development and refinement of digital imaging technology with an on-camera live LCD preview screen has overshadowed SLR's popularity. Nearly all inexpensive compact digital cameras now include an LCD preview screen allowing the photographer to see exactly what the CCD is capturing. However, SLR is still popular in high-end and professional cameras, because the pixel resolution, contrast ratio, and color gamut of an LCD preview screen cannot compete with the clarity and shadow detail of a direct-viewed optical SLR viewfinder.